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The most shade-tolerant species of tree to establish in the process of forest succession. The seedlings of climax species can grow in the shade of the parent trees, ensuring their dominance indefinitely. A disturbance such as fire may kill the climax species, allowing pioneer or earlier successional species to re-establish for a time. White spruce (Picea glauca) is an example of a climax species in the northern forests of North America.